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Renew Checklist of Spiders (Aranei) of Iran



Ghavami Sahra
 
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ABSTRACT

To date 244 species of spiders belonged to 33 families have been recorded from Iran. Sixty two species, 4 genera and 8 families are reported for the first time from Iran in 2002-2006. Spiders were collected from branches, leaves, flowers and trunks of trees and bushes, on the ground, under the stones and grasses by Steiner and Baggiolini method, bottle, aspirator, pitfall trap, pans and insect net. Geolycosa altera Roewer, 955 was found to be junior synonym of Honga radiata (Latreille, 817). Four species of Lycosid spiders were transferred to Trochosa: T. austrocaspia (Roewer, 955), T. flavichelis (Roewer, 955), T. nava (Roewer, 955), T. persia (Roewer, 955).

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  How to cite this article:

Ghavami Sahra , 2006. Renew Checklist of Spiders (Aranei) of Iran. Pakistan Journal of Biological Sciences, 9: 1839-1851.

DOI: 10.3923/pjbs.2006.1839.1851

URL: https://scialert.net/abstract/?doi=pjbs.2006.1839.1851

INTRODUCTION

The first spider collections in persia (Iran) were made by Keyserling and Bienert in 1859. The first published spider record from Iran was by Thorell 1983, who described a new species of Artema. In the first half of 20th century three papers dealt partly with spiders from Iran. Namely black-widows and Argiope have been published: Pocok (1903), Werner (1936) and Pavlosky (1942). In the beginning of the 20th century Russian expenditions to Iran were undertaken by Filipovich in 1904 and by Kirichenko in 1914. Like Keyserling’s material, their collections were stored in ZISP (Zoological Institute, St. Petersburg). While these spiders were partially identified by the late Kharitonov, this data remained unpublished. In the 1960s and 1970s, several expendations to Iran were organized by Austrain, Italian, American, Swiss and German and in 2000s Russian zoologists. Partial results of these expenditions were published in several faunistic and toxonomic papers: Roewer (1955,1959), Brignoli (1972, 1978, 1979, 1980, 1981, 1982), Senglet (1974), Wesolowska (1986), Proszynski (1992), Ovtsharenko et al. (1994), Saaristo and Tanasevitch (1995), Wunderlich (1995), Logunov (1999, 2001a,b), Logunov and Marusik (1999, 2000). Also, Twelve published abstracts and papers by Iranian researchers (Mirshamsi and Darvish, 1997; Goodarzi and Akbari, 1998; Goodarzi et al., 1998; Mozaffarian et al., 1998; Ghavami et al., 2004a, b; Ghavami, 2005, 2006a) and four project research had done by the author. Information on the occurrence and distribution of spiders within Iran is summarized in checklist given blow. New recorded species, genera and families are marked by astrisks(*).

SERVEY OF SPECIES

Agelenidae
Agelena labyrinthica (Clerck, 1757)*
Records: Mazandaran: Salman Shahr and Chaloos, Golestan: Nodijeh, Chardeh, Raamian and Kordkooy, (Ghavami et al., 2004a), Golestan, Mazandaran, (Ghavami et al., 2004a).
Agelena orientalis C.L.Koch,1841
Records: Mazandaran, (Brignoli, 1978).
Cicurina sp.*
Records: Golestan: Gonbad (Ghavami, 2005)

Araneidae
Aculeperia ceropegia (Walckenaer,1802)
Records: Kerman, (Roewer, 1955).
Agalenatea redii (Scopoli, 1763) *
Records: Golestan: Raamian and Gonbad (Ghavami et al., 2004b).
Araniella inconspicua*
Records: Golestan: Raamian (Ghavami et al., 2004a).
Araniella alpica (L.Koch,1869)*
Records: Golestan: Raamian, Nodijeh (Ghavami et al., 2004a).
Araniella cucurbitina (Clerck, 1757)
Records: Golestan: Nodijeh, Zarringol, Tooskaostan (Ghavami et al., 2004b),
Mazandaran (Roewer, 1955; Ghavami, 2006b).
Araniella opistographa (Kulcuzynski, 1905)*
Records: Golestan, (Mozaffarian et al., 2004).
Araneus angulatus Clerck, 1757
Records: Guilan and Mazandaran (Mozaffarian et al., 1998).
Araneus diadematus Clerck,1757 *
Records: Golestan: (Ghavami et al., 2004a), Guilan, Mazandaran (Ghavami, 2005; Ghavami, 2006c).
Argiope bruennichi (Scopoli 1772)
Records: Fars (Roewer, 1955), Zanjan, Golestan: Tooskaostan, Guilan: Roodbar,
Loushan (Ghavami et al., 2004a), Mazandaran, (Mozaffarian et al., 1998; Ghavami, 2005; Ghavami, 2006a).
Argiope clarckii Blackwall,1865
Records: Booshehr (Pocok, 1903; Levy, 1997).
Argiope lobata (Pallas, 1772)
Records: Zanjan (Ghavami et al., 2004b), Ardebil: Moghan (Ghavami, 2005), Fars and Kerman (Roewer, 1955).
Cercidia sp.*
Records: Guilan: Roodbar, Zanjan: Soomeahbar, Vaneaser, Alazain, Jozargh, Zarringol, Hendezamin, Golestan: Tooskaostan, (Ghavami et al., 2004b).
Cyclosa conica (Pallas, 1772)*
Records: Golestan: Kordkooy, Tooskaostan, Zanjan: Hendezamin, (Ghavami et al., 2004a), Mazandaran, (Ghavami, 2006b).
Cyphepeira silvicultrix (C.L.Koch, 1845)
Records: Guilan, (Roewer, 1955).
Hypsosinga pygmaea (Sundevall, 1831)
Records: Guilan (Mozaffarian et al., 1998), Golestan (Ghavami et al., 2004a).
Hypsosinga sanguinea (C.L.Koch, 1845)*
Records: Golestan: Gonbad, Hivechi, (Ghavami et al., 2004a), Mazandaran (Ghavami, 2006c).
Hypsosinga albovittata (Westring, 1851)*
Records: Tehran: Varamin, Jafar Abad, (Ghavami, 2005).
Larinioides cornutus (Clerck, 1758)
Records: Khorram Abad, Mazandaran, Fars, West Azerbayjan and Kerman (Roewer, 1955), Golestan, (Ghavami, 2006b).
Larinioides folium (Schranek, 1803)
Records: Guilan, Mazandaran, (Mozaffarian et al., 1998).
Neoscona adianta (Walckenaer, 1802)
Records: Guilan, Mazandaran (Mozaffarian et al., 1998),
Golestan: Tooskaostan, Alazeman, Beraftan, Nodijeh, Kordkooy, Gonbad, Bailer, Hivechi, Zanghian, Toushan, Hashem Abad, Fazel Abad, (Ghavami et al., 2004a), Ardebil: Moghan, Oltan, (Ghavami, 2005),
Mangora acalypha (Walckenaer, 1802)
Records: Golestan: Tooskaostan, Beraftan, Nazar Abad, Guilan: Roodbar, Ali Abad, (Ghavami et al., 2004b).
Singa licina (Audouin, 1802)
Records: Mazandaran, (Mozaffarian et al., 1998).

Amaurobiidae*
Records: Golestan: Raamian, (Ghavami et al., 2004b).

Atypidae:*
Atypus sp.*
Records: Mazandaran (Goodarzi, 1994)

Clubionidae
Clubiona mildei L. Koch, 1864
Records: Kerman (Roewer, 1955)
Clubiona neglecta O.P. Cambridge, 1862*
Records: Golestan: Kordkooy, Karkandeh, (Ghavami, 2005).
Clubiona phragmitis C.L. Koch 1843
Records: Guilan and Mazandaran, (Mozaffarian et al., 1998).

Cybaeidae
Cybaeus angustiarum L. Koch, 1868
Records: Guilan or Kerman, (Roewer, 1955).

Dictynidae
Achaeodictyna sp.
Records: Esfahan (Mozaffarian and Marusik, 2001)
Argenna patula *
Records: Tehran: Varamin, Jafar Abad, (Ghavami, 2005).
Dictyna latens (Fabricius, 1775) *
Records: Guilan: Roodbar, Golestan: Gonbad, (Ghavami et al., 2004a).
Nigma flavescens (Walckenaer, 1825)*
Records: Kashan, (Ghavami et al., 2004b).

Dysderidae
Dysdera asiatica Nosek, 1905
Records: Esfahan, Kerman (Roewer, 1955).
Dysdera concinna L. Koch, 1878
Records: Iran (Platnik, 2000)
Dysdera erythrina (Walckenaer, 1802)
Records: Esfahan and (Caspian Sea) (Roewer, 1955).
Harpactea parthica Brignoli 1980
Records: Khorasan (Brignoli, 1980).

Eresidae
Eresus sp.*
Records: Guilan: Roodbar, Rostam Abad, Ghazvin: Manjil, (Ghavami et al., 2004b)
Tehran: Varamin, Jafar Abad, (Ghavami, 2005).
Dresserus sp.*
Records: Ghazvin: Tarom, (Ghavami et al., 2004a).
Stegodyphus dufouri (Audouin, 1826)
Records: Province unclear (Roewer, 1955).
Stegodyphus pacificus (Pocok, 1889)
Records: Fars, Kerman (Kraus and Kraus, 1988).

Eusparassidae
Olios xerxes (Pocok, 1889)
Records: Iran, (no locality data) (Platnik, 2000).
Sparassus walckenaeri (Audouin, 1827)
Records: Kerman and Mazandaran, (Roewer, 1955; Levy, 1989).
Micrommata virecens (Clerck, 1757)*
Records: Tehran: Varamin, Jafar Abad, Ardebil: Moghan, Oltan, (Ghavami, 2005).

Filistatidae
Filistata insidiatrix (Forskal, 1775)
Records: Mazandaran (Brignoli, 1982).
Filistata sp.
Records: Roodbar, Zanjan: Gilvan, (Ghavami et al., 2004b).
Zaitunia alexandri Brignoli, 1982
Records: Fars (Brignoli, 1982).
Zaitunia medica Brignoli, 1982
Records: Esfahan (Brignoli, 1982).
Zaitunia persica Brignoli, 1982
Records: Fars (Brignoli, 1982).

Gnaphosidae
Aphantaulax sp.*
Records: Guilan: Roodbar, Golestan: Chardeh, Zanjan: Hendezamin, Ghazvin, (Ghavami et al., 2004a).
Drassodes lapidosus (Walckenaer, 1802)
Records: Guilan, East and probably West Azerbayjan and Kerman (Roewer, 1955).
Drassodes sp.
Records: Fars: Neyriz, Darab, Jahrom, Estahban, (Ghavami et al., 2004a).
Gnaphosa sp.*
Records: Fars: Darab (Ghavami et al., 2004b).
Haplodrassus sp.*
Records: Fars: Darab, Remestagh, (Ghavami, 2005).
Micaria sp.*
Records: Golestan: Gonbad, Tehran: Varamin, Jafar Abad, Goltapeh, Fars: Darab, Remestagh (Ghavami, 2005).
Nomisia exornata (C.L.Koch,1839)
Records: Fars (Roewer, 1955).
Nomisia ripariensis (O.Pickard- Cambridge, 1872)
Records: Mazandaran and Kerman (Roewer, 1955), Tehran, (Mozaffarian et al., 1998).
Phaeocedus braccatus (L. Koch,1878)
Records: Mazandaran (Mozaffarian et al., 1998).
Pterotricha conigera Spassky, 1941
Records: West or East Azerbayjan (Roewer, 1955).
Pterotricha lentiginosa (C.L.Koch,1837)
Records: Fars, East Azerbayjan (Roewer, 1955).
Pterotricha loeffleri (Roewer, 1955)
Records: Tehran, (Roewer, 1955).
Scotophaeus sp.*
Records: Zanjan: Tarom, (Ghavami et al., 2004a).
Synaphosus neali (Ovtsharenko et al., 1994)
Records: Baloochestan (Ovtsharenko et al., 1994).
Synaphosus shirin (Ovtsharenko et al., 1994)
Records: Kermanshah (Ovtsharenko et al., 1994).
Talanites dunini Platnick et Ovtsharenko, 1994
Records: Guilan (Mozaffarian et al., 1998).
Zelotes femellus (C.L. Koch, 1866)
Records: East Azerbayjan (Roewer, 1955).
Zelotes longipes (C.L. Koch, 1866)
Records: Guilan (Mozaffarian et al., 1998).
Zelotes sp.
Records: Golestan (Ghavami et al., 2004b), Mazandaran, (Ghavami, 2005; Ghavami, 2006b).

Hahniidae
Antistea sp.*
Records: Fars: Darab (Ghavami, 2005).

Linyphiidae
Erigone dentipalpis (Wider, 1834)
Records: Kerman or Guilan, (Roewer, 1955).
Frontinella frutetorum (C.L.Koch, 1834)*
Records: Tehran, (Goodarzi, 1994), Guilan: Roodbar, Golestan: Nodijeh, Raamian, Zanjan: mamalan, Soomeahbar, Hendezamin, Vaneser, Tarom (Ghavami et al., 2004b)
, Golestan, Mazandaran, (Ghavami, 2006c).
Gnathonarium dentatum (Wider, 1834)
Records: Kerman or Guilan (Roewer, 1955).
Lepthyphantes zimmermanni Bertkau, 1890*
Records: Tehran: Varamin, Goltapeh (Ghavami, 2005).
Lepthyphantes iranicus Saaristo and Tanasevitch, 1995
Records: Mazandaran (Saaristo and Tanasevitch, 1995).
Lepthyphantes nebulosus (Sundevall, 1830)
Records: Kerman (Roewer, 1955).
Lepthyphantes sbordonii Brignoli, 1970
Records: Mazandaran (Brignoli, 1970).
Leptorhoptrum robustum (Westring, 1851)*
Records: Guilan, Zanjan, Tarom (Ghavami et al., 2004a).
Linyphia triangularis (Clerck, 1758)
Records: Kerman or Guilan (Roewer, 1955).
Linyphia sp.
Records: Zanjan: Tarom, Ghazvin, Tarom (Ghavami et al., 2004a), Tehran: Varamin, Goltapeh, Golestan, (Ghavami, 2005; Ghavami, 2006b).
Microlinyphia sp.*
Records: Golestan: Tooskaostan, Zanjan: Vaneser, Guilangasheh (Ghavami et al., 2004b), Tehran: Varamin, Goltapeh, Jafar Abad, (Ghavami, 2005)
Golestan, Mazandaran, (Ghavami, 2006c).
Oedothorax apicatus (Blackwall, 1850)*
Records: Khorasan (Mozaffarian et al., 2004).
Prinerigone vagans (Audouin, 1826)
Records: Guilan and Mazandaran (Mozaffarian et al., 1998).
Tenuiphantes perseus (Van Helsdingen et al., 1977)
Records: Guilan (Van Helsdingen et al., 1977).

Loxoscelidae*
Loxosceles sp.*
Records: Tehran (Goodarzi, 1994)

Lycosidae
Acantholycosa sp.*
Records: Golestan (Goodarzi, 1994)
Allohogna singoriensis (Laxmann, 1770)
Records: East Azerbayjan, (Roewer, 1955).
Alopecosa pulverulenta (Clerck, 1757)*
Records: Golestan, Tehran: Varamin, Jafar Abad, (Ghavami, 2005).
Arctosa cinerea (Fabricius, 1775)
Records: East and probably West Azerbayjan, (Roewer, 1955).
Arctosa leopardus (Sundevall, 1833)
Records: Guilan and Mazandaran, (Mozaffarian et al., 1998).
Arctosa tbilisiensis Mcheidze, 1947*
Records: Eilam (Mozaffarian et al., 2004).
Aulonia albimana (Walckenaer, 1805)*
Records: Fars: Darab, (Ghavami, 2005).
Evippa fortis Roewer, 1955
Records: Kerman, (Roewer, 1955).
Evippa luteipalpis Roewer, 1955
Records: Kerman (Roewer, 1955).
Evippa praelongipes (O.Pickard-Cambridge, 1870)
Records: Mazandaran (Roewer, 1955).
Geolycosa(?) atroscopulata Roewer, 1955
Records: Kerman (Roewer, 1955).
Honga radiata (Latreille, 1817)
Records: Mazandaran (Roewer, 1955).
Lycosa suzukii Yaginuma, 1973*
Records: Tehran (Goodarzi, 1994)
Lycorma(?) nigrichelis Roewer, 1955
Records: Mazandaran and East Azerbayjan, (Roewer, 1955).
Pardosa agrestis (Westring, 1861)*
Records: Golestan: Fazel Abad, Gharan Abad, Tooskaostan, Guilan: Ali Abad (Ghavami et al., 2004b; Ghavami, 2006c), Golestan, Ardebil: Moghan (Ghavami, 2005).
Pardosa agricola (Thorell, 1856)
Records: Kerman, Tehran, Markazi and East Azerbayjan, (Roewer, 1955).
Pardosa amentata (Clerck,1757) *
Records: Golestan: Tooskaostan,
Gonbad, Fars: Neyriz, (Ghavami, 2004a, 2005).
Pardosa condolens (O.Pickard-Cambridge, 1885)
Records: Esfahan (Roewer, 1955).
Pardosa donabila Roewer, 1955
Records: Fars, West or East Azerbayjan, Tehran and Mazandaran (Roewer, 1955).
Pardosa lugubris (Walckenaer, 1802)
Records: Kerman, (Roewer, 1955).
Pardosa monticola (Clerck, 1757)*
Records: Golestan: Fazel Abad, Hashem Abad, (Ghavami, 2005).
Pardosa palustris (Linnaeus, 1758)
Records: Golestan (Goodarzi, 1994)
Pirata piraticus (Clerck, 1758)
Records: West or East Azerbayjan, (Roewer, 1955).
Trochosa austrocaspia (Roewer, 1955)
Records: Mazandaran (Roewer, 1955).
Trochosa flavichelis (Roewer, 1955)
Records: Mazandaran, Guilan or Kerman (Roewer, 1955).
Trochosa nava (Roewer, 1955)
Records: Kerman, (Roewer, 1955).
Trochosa persica (Roewer, 1955)
Records: Guilan or Kerman (Roewer, 1955).
Trochosa urbana (O.Pickard-Cambridge, 1876)
Records: Fars, Mazandaran and Kerman (Roewer, 1955).
“Trochosa” impercussa Roewer, 1955
Records: Kerman (Roewer, 1955).
“ Trochosa”loeffleri Roewer, 1955
Records: Esfahan (Roewer, 1955).
Trochosa terricola Thorell, 1856
Records: Kerman (Roewer, 1955).
Wadicosa sp.
Records: Eilam and Khoozestan (Mozaffarian et al., 1998).

Metidae*
Zygiella x-notata (Clerck, 1757) *
Records: Golestan, (Ghavami et al., 2004a).

Miturgidae*
Cheiracanthium erraticum (Walckenaer, 1802)*
Records: Tehran: Varamin, Jafar Abad, Goltapeh, Ardebil: Moghan (Ghavami, 2005).
Cheiracanthium pennyi O.P.-Cambridge, 1873*
Records: Golestan: Gonbad, Bailer, Hivechi, Ardebil: Moghan (Ghavami, 2005).
Cheiracanthium mildei C.L.Koch, 1864
Records: Kerman (Roewer, 1955), Tehran (Goodarzi, 1994), Tehran: Varamin, Jafar Abad, Pishva (Ghavami, 2005).

Nemestidae
Raveniola niedermeyeri (Brignoli, 1972)
Records: Golestan, (Brignoli, 1972).

Oecobiidae
Oecoblus sp.*
Records: Khorasan: Kashmir, Tehran: Varamin, Jafar Abad, Golestan: Gonbad, Bailer (Ghavami, 2005).
Uroctea durandi (Latreille, 1809)
Records: Kerman (Roewer, 1955).

Oxyopidae
Oxyopes heterophthalmus Latreille, 1804*
Records: Golestan (Mozaffarian et al., 2004).
Oxyopes lineatus (Latreille, 1806)
Records: East or West Azerbayjan, (Roewer, 1955), Tehran, (Goodarzi, 1994), Golestan: Beraftan, (Ghavami, 2004b).
Peucetia sp.*
Records: Fars: Darab (Ghavami, 2005).

Palpimanidae
Palpimanus gibbulus Dufour, 1820
Records: West Azerbyjan, Yazd (Roewer, 1955).

Philodromidae
Philodromus rufus (Walckenaer, 1825)*
Records: East Azerbayjan (Mozaffarian and Khalil, 1999),
Golestan: Tooskaostan, Zarringol, Kordkooy, Beraftan, Zanjan, Guilan, (Ghavami et al., 2004a).
Tibellus oblongus (Walckenaer, 1802)
Records: Guilan, Mazandaran, Golestan: Gonbad, Hivechi (Ghavami, 2004a, 2005; Ghavami, 2006b).
Philodromus cespitum (Walckenaer, 1802)*
Records: Tehran: Varamin, Jafar Abad, Ardebil: Moghan (Ghavami, 2005).
Philodromus dispar Walckenaer, 1825
Records: (Caspian Sea), (Roewer, 1955).
Philodromus margaritatus (Clerck, 1758)
Records: Mazandaran or Kerman, (Roewer, 1955).
Thanatus vulgaris
Records: Tehran (Goodarzi and Akbari, 1998), Eilam, Tehran, Khorasan, (Mozaffarian and Marusik, 2001).
Thanatus formicinus (Clerck, 1757)*
Records: Tehran: Varamin, Jafar Abad, Pishva, Taghi Abad, Golestan: Gonbad, Fars: Darab, Khorasan: Kashmar (Ghavami, 2005).
Thanatus striatus C.L.Koch,1845*
Records: Tehran: Varamin, Jafar Abad, Fars: Darab (Ghavami, 2005).

Pholcidae
Artema doriai Thorell in Simon, 1893
Records: Kerman, Mazandaran and Tehran (Roewer, 1955).
Psilochorus simoni (Berland 1911) *
Records: Golestan: Tooskaostan, Kordkooy, Chardeh, Bandar Gaz,
Ghazvin (Ghavami et al., 2004a).
Pholcus phalangioides (Fuesslin, 1775) *
Records: Golestan: Chardeh, Bandar Gaz, Guilan: Roodbar, (Ghavami et al., 2004b).
Pholcus alticeps Spassky, 1932
Records: Guilan, Golestan, Mazandaran, (Senglet, 1974).
Pholcus armeniacus Senglet, 1974
Records: West Azerbayjan (Senglet, 1974).
Pholcus hyrcanus Senglet, 1974
Records: Golestan (Senglet, 1974).
Pholcus medicus Senglet, 1974
Records: Zanjan, Markazi (Senglet, 1974).
Pholcus opilionoides (Schrank, 1781)?
Records: Kerman (Roewer, 1955).
Pholcus persicus Senglet, 1974
Records: Fars (Senglet, 1974).

Pisauridae
Pisaura mirabilis (Clerck, 1757)
Records: Mazandaran (Mozaffarian et al., 1998),
Ardebil: Moghan, Oltan, Majid Abad (Ghavami, 2005).

Salticidae
Aelurillus iranus (Roewer, 1955)
Records: Kerman (Roewer, 1955, Proszynski, 1966).
Aelurillus concolor Kulezynski, 1901
Records:Sabzevar (Roewer, 1955).
Ballus depressus (Walckenaer, 1802)
Records: Mazandaran (Logunov and Marusik, 2000).
Bianor albimaculatus (Lucas, 1846)
Records: Guilan and Mazandaran (Mozaffarian et al. 1998; Logunov, 2001a), Fars, Khoozestan (Logunov, 2001b), Zanjan: Tarom (Ghavami et al., 2004a).
Bianor aurocinctus (Ohlert, 1865)
Records: Khorram Abad (Roewer, 1955).
Chalcoscirtus (Chalcoscirtus) iranicus (Logunov and Marusik, 1999)
Records: Tehran.
Chalcoscirtus (s.str.) infimus
Records: Tehran, (Logunov and Marusik, 1999, 2000).
Chalcoscirtus (s.str.) lepidus Wesolowska, 1986
Records: Tehran (Logunov et al., 2001).
Chalcoscirtus (s.str.) nigritus (Thorell, 1875)
Records: Fars, East Azerbayjan, (Logunov and Marusik, 1999).
Chalcoscirtus (s.str.) hosseinieorum sp.n.
Records: Fars (Logunov et al., 2001).
Chalcoscirtus (Chalcoscirtus) parvulus
Records: Tehran (Logunov et al., 2001).
Chinnatus caucasicus (Logunov, 1999)
Records: Mazandaran (Roewer, 1955), Khorram Abad, (Logunov, 1999).
Cyrba algerina (Lucas, 1846)
Records: Fars, Tehran (Logunov et al., 2001).
Cyrba ocellata (Kroneberg, 1875)
Records: Tehran (Mozaffarian et al., 1998), Mazandaran (Logunov et al., 2001).
Euophrys frontalis (Walckenaer, 1802)
Records: Mazandaran (Mozaffarian, 2000), Fars: Estahban (Ghavami, 2005).
Evarcha arcuata (Clerck, 1758)
Records: Mazandaran
Evarcha negevensis Proszynski, 1992
Records: Tehran (Logunov et al., 2001).
Evarcha praecincta (Simon, 1893)
Records: Fars (Logunov et al., 2001).
Heliophanus (s.str.) cupreus (Walckenaer, 1802)
Records: East Azerbayjan, Mazandaran, (Wesolowska, 1986; Marusik et al., 2000), Zanjan, Tehran: Varamin, Jafar Abad (Ghavami, 2005).
Heliophanus (s.str.) decoratus C.L. Koch, 1875
Records: Fars (Logunov et al., 2001).
Heliophanus (s.str.) dunini Rakov and Logunov, 1997
Records: Tehran (Marusik et al., 2000).
Heliophanus (s.str.) equester L. Koch, 1867
Records: West Azerbayjan, Tehran and Fars (Logunov et al., 2001).
Heliophanus (s.str.) flavipes Huhn, 1831
Records: East Azerbayjan, Mazandaran, Tehran (Wesolowska, 1986), Mazandaran, Tehran (Logunov et al., 2001), Golestan: Gonbad, Kordkooy, Tehran: Varamin, Jafar Abad, (Ghavami, 2004b, 2005),
Heliophanus forcipifer Kulezynski, 1895
Records: Yazd (Mozaffarian and Marusik, 2001)
Heliophanus fulgens (O.Pickard-Cambridge, 1876)
Records: Yazd, (Mozaffarian et al., 1998) Esfahan, Fars, (Logunov et al., 2001).
Heliophanus (Heliophanus) edentulus
Records: Tehran (Logunov et al., 2001).
Heliophanus (s.str.) glaucus Bosenberg and Lenz, 1895
Records: Fars (Logunov et al., 2001).
Heliophanus (s.str.) iranus Wesolowska, 1986
Records: Tehran (Wesolowska, 1986).
Heliophanus ignorabilis Wesolowska, 1986
Records: Mazandaran (Wesolowska, 1986).
Heliophanus ledeufiilus (S.str.Hahn,1831)*
Records: Golestan: Tooskaostan (Ghavami et al., 2004b).
Heliophanus (s.str.) mordax (O.Pickard-Cambridge, 1872)
Records: Tehran, Fars, Mazandaran (Marusik et al., 2000).
Heliophanus (s.str.) verus Wesolowska, 1986
Records: Mazandaran (Wesolowska, 1986), Kerman, (Logunov and Marusik, 2000).
“Hyllus” insularis,
Records: Yazd, Mazandaran (Logunov, 2001a), Zanjan, Guilan (Ghavami et al., 2004a).
Iannattus rectangularis Proszynski, 1992
Records: Baloochestan (Proszynski, 1992).
Langona redii (Savigny and Audouin, 1825).
Records: Fars (Gettmer, 1983)
Macaroeris nidicolens (Walckenaer, 1802)
Records: Golestan (Logunov and Marusik, 2000).
Mendoza canestrinii (Ninni in Canestrini and Pavesi, 1868).
Records: Guilan, Golestan and Mazandaran (Mozaffarian et al., 1998).
Menemerus semilimbatus (Hahn, 1827)
Records: Mazandaran (Logunov et al., 2001).
Mexcala farsensis (Logunov, 2001a)
Records: Fars (Logunov, 2001b).
Mogrus neglectus
Records: Tehran and Fars (Logunov et al., 2001).
Myrmarachne tristis
Records: Khoozestan (Marusik, 1996).
Neon (Dicroneon) levis
Records: Tehran, (Logunov and Marusik, 2000).
Neon (Dicroneon) rayi
Records: Tehran (Logunov et al., 2001).
Neon (s.str.) reticulatus (Blackwall, 1853)
Records: East Azerbayjan, Mazandaran (Logunov et al., 2001).
Pellenes (Pelmultus) brevis
Records: Tehran (Logunov et al., 2001).
Pellenes (Pelmultus) epularis (O.Pickard-Camb-ridge, 1872)
Records: Fars (Marusik et al., 2000), Tehran (Logunov et al., 2001).
Pellenes (Pellenes) seriatus (Thorell, 1875)
Records: Golestan and Kermanshah (Logunov and Marusik, 2000).
Philaeus chrysops (Poda, 1761)
Records: Tehran, Golestan and Mazandaran (Logunov et al., 2001), Eilam (Mozaffarian et al., 2004).
Phintella castriesiana (Grube, 1861)
Records: Mazandaran
Khorram Abad and Mazandaran (Logunov et al., 2001).
Phlegra fascita (Hahn, 1826)
Records: Mazandaran (Logunov et al., 2001).
Phlegra tetralineata (Caporiacco, 1939)
Records: Tehran (Azerkina and Logunov, 2001).
Plexippoides flavescens (O.Pickard- Cambridge, 1872)
Records: Kerman (Kuznetsov, 1929), Mazandaran (Roewer, 1955), Tehran (Mozaffarian et al., 1998).
Plexippoides coccineus
Records: Tehran (Logunov et al., 2001).
Pseudicius rudakii Proszynski, 1992
Records: Fars (Proszynski, 1992).
Pseudeuophrys erratica (Walckenaer, 1826)
Records: Mazandaran (Logunov et al., 2001).
Pseudicius palaestinensis Strand, 1915
Records: (Logunov et al., 2001), Fars, Yazd (Mozaffarian and Marusik, 2001), Tehran.
Pseudicius rudakii Proszynski, 1992
Records: Fars, (Proszynski, 1992).
Pseudicius spasskyi
Records: Guilan (Wesolowska, 1986).
Salticus cingulatus (Panzer, 1797)
Records: Kerman, (Roewer,1955).
Salticus scenicus (Clerck, 1757)*
Records: Zanjan (Ghavami et al., 2004a), Khorasan: Kashmar (Ghavami, 2005).
Salticus tricinctus (C.L.Koch, 1846)
Records: Tehran, Mazandaran and Fars.
Sibianor aurocinctus (Ohlert, 1865)
Records: Mazandaran (Roewer, 1955).
Sitticus ammophilus (Thorell, 1875)
Records: Iran, (no locality data) (Wesolowska, 1986; Logunov and Marusik, 2000).
Sitticus relictarius Logunov, 1998
Records: Mazandaran
Stenaelurillus marusiki Logunov et al. (2001)
Records: Fars (Logunov, 2001b).
Synageles dalmaticus (Keyserling, 1863)
Records: Yazd, Markazi province, Saveh (Mozaffarian et al., 1998), Tehran (Logunov et al., 2001).
Talavera aequipes (O.Pickard- Cambridge,1872)
Records: Mazandaran (Logunov et al., 2001).
Thyene imperialis (Rossi, 1846)
Records: Zanjan: Markazi province: Saveh (Mozaffarian et al., 1998), Zanjan: Tashvir, Tarom, Jozargh, Golestan: Chardeh, Zarringol, Azad Shahr, Gonbad, Hivechi, Hashem Abad, Tooskaostan, Guilan, Mazandaran (Ghavami, 2004b, 2005; 2006c), Tehran: Varamin, Jafar Abad, Goltapeh, Taghi Abad, Fars: Darab (Ghavami, 2005).

Scytodidae
Scytodes fusca (Walckenaer, 1837)
Records: Kerman (Roewer, 1955).
Scytodes thoracica (Latreille, 1804)
Records: Caspian Sea (Roewer, 1955) Zanjan, Golestan (Ghavami et al., 2004a).

Tetragnathidae
Pachygnatha degeeri Sundevall, 1830
Records: Kerman, West or East Azerbayjan (Roewer, 1955).
Tetragnatha extensa (Linnaeus, 1758)
Records: Mazandaran, West Azerbayjan (Roewer, 1955), Guilan: Roodbar, Golestan: Raamian, Fazel Abad, Bandar Gaz, Zanghian, Gonbad, Hashem Abad, Mazandaran (Ghavami, 2004a, 2005, 2006a).
Tetragnatha javana (Thorell, 1890) *
Records: Guilan: Roodbar, Golestan: Toshan, Kordkooy, Karkandeh, Gonbad, Hashem Abad, Mazandaran (Ghavami, 2004a, 2005; 2006b).
Tetragnatha montana
Records: Golestan: Gonbad, Toshan, Hashem Abad, Raamian, Mazandaran (Ghavami, 2004a, 2005; 2006c).
Tetragnatha nigrita Lendl, 1886*
Records: Golestan, Mazandaran (Ghavami, 2006b).

Theridiidae
Anelosimus aulicus (C.L.Koch,1838)*
Records: Guilan, Zanjan (Ghavami et al., 2004a).
Dipoena prona (Menge, 1868)*
Records: Zanjan: Tarom, (Ghavami et al., 2004a).
Enoplognatha mandibularis (Lucas, 1846)
Records: Mazandaran (Roewer, 1955).
Enoplognatha mediterranea
Records: Ardebil (Ghavami, 2004a).
Latrodectus dahli Levi, 1959
Records: Iran, (no locality data) (Platnik, 2000).
Latrodectus geometricus C.L.Koch, 1841
Records: Khorasan (Rafiinejad et al., 2000).
Latrodectus palidus O. Pickard- Cambridge, 1872
Records: Khorasan (Mirshamsi and Darvish, 1997; Rafiinejad et al., 2000).
Latrodectus tredecimguttatus (Rossi, 1790)
Records: Iran, (no locality data) (Werner, 1936; Goodarzi et al., 1998), Hormozgan, Booshehr, Mazandaran (Roewer, 1955), Khorasan (Mirshamsi and Darvish, 1997; Rafiinejad et al., 2000), Golestan: Hashem Abad (Ghavami et al., 2004b).
Robertus lividus (Blackwall, 1836)
Records: Esfahan, (Roewer, 1955).
Steatoda albomaculata (Degeer,1778)*
Records: Zanjan: Tarom, Goocan, Tashvir, Hendezamin, Gilvan, Alzain, Guilan: Roodbar, Ghazvin, Mazandaran, (Ghavami et al., 2004a; Ghavami, 2006a).
Steatoda paykulliana (Walckenaer, 1806)*
Records: Eilam (Mozaffarian et al., 2004), Golestan: Gonbad, Bailer, Hashem Abad, Ardebil: Moghan, Goosloo, Aghdam, Oltan (Ghavami, 2005).
Steatoda phalerata (Panzer, 1801)
Records: Mazandaran (Roewer, 1955).
Theridion hemerobius
Records: Guilan, Mazandaran (Mozaffarian et al., 1998, 2004).
Theridion melanurum Hahn, 1831
Records: Zanjan (Ghavami, 2006c)
Theridion tinctum (Walckenaer, 1802)
Records: Kerman or Guilan, East Azerbayjan (Roewer, 1955)
Theridion varians Hahn, 1831
Records: Fars, Kerman, (Roewer, 1955).
Theridion impressum C.L.Koch,1881*
Records: Golestan: Gonbad, Bailer, Ghareghaj, Tehran: Varamin, Taghi Abad, Ardebil: Moghan, Aghdam (Ghavami, 2005).
Theridion simile, C.L.Koch,1836*
Records: Golestan: Kordkooy, Guilan, Mazandaran (Ghavami et al., 2004a; Ghavami, 2006b).

Thomisidae
Diaea dorsata (Fabricius, 1777)*
Records: Tehran: Varamin, Jafar Abad (Ghavami, 2005).
Misumena vatia (Clerck, 1757)*
Records: Guilan: Roodbar, Golestan: Soomeaehbar, Raamian, Fazel Abad, Bandar Gaz, Gonbad, Zanjan, Ghazvin, Mazandaran (Ghavami, 2004a, 2005, 2006b), Tehran: Varamin, Jafar Abad (Ghavami, 2005).
Misumenops tricuspidatus (Fabricius, 1775)
Records: Mazandaran (Mozaffarian et al., 1998).
Monaesis acicullus
Records: Ardebil: Moghan, Oltan (Ghavami, 2005).
Runcinia lateralis C.L.Koch,1838*
Records: Tehran (Mozaffarian et al., 2004)
Runcinia sp.
Records: Golestan, Mazandaran (Ghavami, 2004a, 2005; Ghavami, 2006b).
Ozyptila aculipalpa Wunderlich, 1995
Records: Khorram Abad (Wunderlich, 1995).
Ozyptila simplex (O. Pickard- Cambridge, 1862)
Records: Guilan (Mozaffarian et al., 1998, 2004).
Ozyptila grisea Roewer, 1955
Records: Kerman (Roewer, 1955).
Ozyptila rauda
Records: Guilan, Mazandaran (Roewer, 1955).
Ozyptila praticola (C.L.Koch,1837)
Records: Kerman (Roewer, 1955).
Ozyptila sp.
Records: Guilan (Ghavami et al., 2004b).
Synaema globosum (Fabricius, 1775)
Records: Guilan: Roodbar, Ali Abad, Mazandaran (Roewer, 1955), Golestan: Beraftan, Tooskaostan, Toushan, Karkandeh, Kordkooy, Raamian, Zarringol, Moodestan, Zanjan, Mazandaran (Ghavami et al., 2004a, 2005; Ghavami, 2006b).
Thomisus onustus, Walckenaer, 1806
Records: Guilan or Mazandaran (Roewer, 1955), Zanjan: Jozargh, Golestan: Soomeaehbar, Toushan, Kordkooy, Mazandaran (Ghavami, 2004b, 2005, 2006c), Tehran: Varamin, Jafar Abad, Fars: Darab, Ardebil: Moghan (Ghavami, 2005).
Thomisus zyuzini Marusik and Logunov, 1990*
Records: Esfahan, Khorasan (Mozaffarian et al., 2004)
Temarus sp.*
Records: Tehran: Varamin, Jafar Abad (Ghavami, 2004a, 2005).
Xysticus cristatus (Clerck, 1757)*
Records: Golestan: Raamian, Nodijeh, Karkandeh, Kordkooy, Toshan, Zanjan: Guilangasheh, Guilan: Roodbar, Ali Abad, Mazandaran (Ghavami, 2004a, 2005; 2006b), Ardebil: Moghan (Ghavami, 2005).
Xysticus kochi Thorell, 1872*
Records: Guilan: Roodbar, Golestan: Hashem Abad, Tooskaostan (Ghavami et al., 2004a).
Xysticus audax (Schrank, 1803)
Records: Kerman (Roewer, 1955).
Xysticus lanio C.L.Koch,1824
Records: Guilan, Mazandaran (Roewer, 1955), Fars: Darab (Ghavami, 2005).
Xysticus loeffleri Roewer, 1955
Records: Guilan or Mazandaran (Roewer, 1955).
Xysticus tristrami (O. Pickard- Cambridge, 1872)
Records: Fars (Roewer, 1955).
Xysticus luctousus (Blackwall,1836)*
Records: Fars (Roewer, 1955), Ardebil: Moghan, Fars: Estahban, Neyriz (Ghavami, 2005).

Titanoecidae
Nurscia albomaculata (Lucas,1846)*
Records: Tehran (Mozaffarian and Marusik, 2001).
Titanoeca turkmenia Wunderlich, 1995
Records: Mazandaran, (Wunderlich, 1995).
Titanoeca sp.
Records: Khorasan (Mozaffarian et al., 2004).

Uloboridae
Octonoba yesoensis (S.Saito, 1934)
Records: Mazandaran (Brignoli, 1979).
Uloborus plumipes Lucas, 1846
Records: Fars (Brignoli, 1979).
Uloborus walckenaevius Latreille, 1806*
Records: Tehran, Khorram Abad (Goodarzi, 1994)

Zodariidae*
Records: Tehran, Eilam (Mozaffarian et al., 2004).

DISCUSSION

Study of Spiders is young in Iran. Therefore, the most species of spiders haven’t been identified. Two hundred and forty four species of spiders have recorded since 1859 to now. The most of species were reported belonged to Salticidae, Lycosidae, Araneidae and Thomisidae families (70, 30, 21 and 20 species, respectively) and the fewest species belonged to Atypidae, Cybaeidae, Metidae, Nemestidae, Palpimanidae. Pisauridae and Titanoecidae families (Table 1). Spiders were collected from gardens, fields, forests, plains and houses. Some species were collected on different crops (apple, cereal, citrus, cotton, olive, pomegranate and rice) (Table 2).

Table 1: Number of species, new species and new genera of spiders were recorded of Iran
*Families marked by asterisks are reported for the first time from Iran

Table 2: Spider species were collected on different crops
(+) Indicating the spider species were collected on different crops, (-) Indicating the spider species not found

They had high population in gardens and fields and they had important role in pest control. The number of species were similar to spider fauna in Azerbayjan, Turkmenistan, Armenia, Kazakhstan, Mediterranean and the Near East But, spiders identification in these area and Iran is necessary to more study.

ACKNOWLEDGMENTS

I wish to thank to Mr. Bahramishad, Khani and Beirami for their field assistance.

REFERENCES
1:  Azerkina, G.N. and D.V. Logunov, 2001. Separation and distribution of Xysticus cristatus (Clerck, 1758) and X. udax (Schrank, 1803) in Eastern Eurasia, with description of new species from the mountains of central Asia. Arthropoda Selecta, 9: 133-150.

2:  Brignoli, P.M., 1970. Identification Leptyphantes cavernicolo from Iran (Araneae, Linyphidae). Fragment Entomol., 7: 55-60.

3:  Brignoli, P.M., 1972. A new Brachytele from Iran. Rev. Swiss J. Zool., 79: 461-541.

4:  Brignoli, P.M., 1978. Introduce a spider species living in concave. Rev. Swiss J. Zool., 85: 461-541.

5:  Brignoli, P.M., 1979. Identification uloboridae family from palearctic region. Rev. Arachnol., 2: 275-282.

6:  Brignoli, P.M., 1980. A new Harpactea from Iran (Araneae, Dysderidae). Bull. Bri. Arachnol. Soc., 5: 95-97.

7:  Brignoli, P.M., 1981. Studies on the pholcidae (Araneae), 1. Notes on the genera Artema and Physocyclus (Araneae). Bull. Am. Musiem Natl. History, 170: 90-100.

8:  Brignoli, P.M., 1982. Identification filistatidae (Araneae) palearctic. Rev. Arachnol., 4: 65-75.

9:  Goodarzi, H.R., 1994. An introduction to the identification and classification of Iranian spiders. M.Sc. Thesis, Plant Pest and Diseese Research Institute, Tehran Iran.

10:  Goodarzi, H.R. and A. Akbari, 1998. Introducing two sub-families and genera of family Thomisidae. Proceeding of 13th Iranian Plant Protection Congress, IPPC'1998, Iran, pp: 232-232.

11:  Goodarzi, H.R. and A. Akbari, H. Vatanpoor, and S.M. Tabatabii, 1998. Extraction of the Iranian black widow spiders venom collected from Khorasan province and estimating their lethal dose (LD50) (Theridiidae, Laterodectus tredecimgutatus). Proceeding of 13th Iranian Plant Protection Congress, IPPC'1998, Iran, pp: 233-233.

12:  Ghavami, S., M. Ahmadi Damghan, S. Ghannad Amooz, S. Soodi and S. Javadi, 2004. Investigation spider fauna in olive orchards in Northern part of Iran. Proceedings of the 16th Congress on Iranian Plant Protection, CIPP'2004, Tabriz, pp: 463-464.

13:  Ghavami, S., 2004. Role of spiders in biological control in Iran. Sonboleh J. Agric. Sci., 135: 23-24.

14:  Ghavami, S., 2004. Black widow and died bite for human. Sonboleh J. Agric. Sci., 137: 34-34.

15:  Ghavami, S., M. Taghizadeh, G.A. Amin, Z. Karimian and M. Hossieni, 2004. Investigation spider fauna in cotton field of Iran. Plant Pests and Diseases Research Institute. Final Report Project, pp: 45.

16:  Ghavami, S., 2005. Palpita unionalis, the most important olive pest in Iran. Zeiyton J. Agric. Sci., 72: 10-12.

17:  Ghavami, S., 2006. Investigation spider fauna in South of the Caspian Sea. Plant Pests and Disease Research Institute Project.

18:  Ghavami, S., 2006. Abundance of spiders (Arachnida: Araneae) in olive orchards in Northern part of Iran. Pak. J. Biol. Sci., 9: 795-799.
CrossRef  |  Direct Link  |  

19:  Ghavami, S., 2006. Role of spiders in citrus pests control. Sonboleh J. Agric. Sci., 145: 23-23.

20:  Van Helsdingen, P.J., K., Thaler and C. Deltshev, 1977. The tenuis group of Lepthyphantes menge (Araneae, Linyphiidae). Tijdschr. Voor Entomol., 120: 1-54.
Direct Link  |  

21:  Kraus, O. and M. Karus, 1988. The genus Stegodyphus (Arachnida, Araneae). Sibling species, species groups and parallel origin of social living. Verh. Naturwiss. Ver. Hamburg, 30: 151-254.

22:  Levy, G., 1989. The family of huntsman spiders in Israel with annotations on species of the Middle East (Aranae: Sparassidae). J. Zool., 217: 127-176.

23:  Levy, G., 1997. Twelve genera of orb-weaver spiders (Araneae: Araneidae) from Israel. Israel J. Zool., 43: 311-365.

24:  Logunov, D.V., 1998. New species and new records of palaearctic Sitticus (Araneae: Salticidae). Zoosystematica Rossica, 7: 77-83.

25:  Logunov, D.V., 1999. Redefinition of the genus Habrocestoides proszynski, 1992, with establishment of new genus, Chinnatus gen.n. (Araneae: Salticidae). Bull. Bri. Arachnol. Soc., 11: 139-149.

26:  Logunov, D.V. and Y. Marusik, 1999. A brief review of the genus Chalcoscirtus bertkau, 1880 in the fauna of Central Asia and the caucasus (Araneae: Salticidae). Arthropoda Selecta, 7: 263-292.

27:  Logunov, D.V. and Y. Marusik, 2000. Miscellaneous notes on palaearctic salticidae (Arachnida: Aranei). Arthropoda Selecta, 8: 263-292.

28:  Logunov, D.V., 2001. A redefinition of the genera Bianor Peckham and Peckham, 1885 and Harmochirus Simon, 1885, with the establishment of a new genus Sibianor gen.n. (Araneae: Salticidae). Arthropoda Selecta, 9: 221-286.

29:  Logunov, D.V., 2001. New and poorly known species of the jumping spiders (Araneae: Salticidae) from Afghanistan, Iran and Crete. Arthropoda Selecta, 10: 59-66.

30:  Logunov, D.V., Y. Marusik and F. Mozaffarian, 2001. Faunistic review of the jumping spiders of Iran (Aranei: Salticidae). Arthropoda Selecta, 10: 155-167.

31:  Marusik, Y., D.V. Logunov and S. Koponen, 2000. Spiders of Tuva, South Siberia. IBPN FEB RAS., Magadan.

32:  Mirshamsi, O. and J. Darvish, 1997. Identification and study of widow spiders (Genus: Laterodectus) in Khorasan Province. Proceedings of 1st Iranian Congress of Zoology, September 17-18, 1997, Tehran, Iran, pp: 106-.

33:  Mozaffarian, F., H. Bayat Asady and S. Tirgari, 1998. Studies on the abundance of spiders in rice fields in central North Provinces of Iran. Proceeding of 13th Iranian Plant Protection Congress, IPPC'1998, Iran, pp: 48-48.

34:  Mozaffarian, F. and A.A. Khalil, 1999. Investigations on the fauna of spiders apple orchards in Tehran and West Azerbayjan. Proceedings of 14th Iranian Plant Protection Congress, September 5-8, 1999, Isfahan, Iran, pp: 283-.

35:  Mozaffarian, F. and Y.M. Marusik, 2001. A checklist of Iranian spiders. Arthropoda Selecta, 10: 67-74.

36:  Mozaffarian, F., H. Roodsari and M.T. Mobasheri, 2004. Investigation spider fauna in cereal fields in Iran. Proceedings of the 16th Congress on Iranian Plant Protection, (CIPP'04), Tabriz, pp: 462-463.

37:  Ovtsharenko, V.I., G. Levy and N.I. Platnik, 1994. A review of the ground spider genus Synaphosus (Araneae: Gnaphosidae). Am. Mus. Novit., 3095: 1-27.
Direct Link  |  

38:  Pavlosky, E.N., 1942. Venomous animals of Central Asia and Iran. Tadzh. Fil. AN SSSR., Stalinabad, pp: 116.

39:  Platnik, N.I., 2000. The world spider catalog. American Museum of Natural History, pp: 81-87. http://research.Amnh.Org/entomology/spiders/catalog.

40:  Pocok, R.I., 1889. Arachnida, chilopoda and crustacea. aitchison, the zoology of the Afghan delimitation commission. Trans. Linn. Soc. London. Ser., 2: 110-121.

41:  Pocok, R.I., 1903. Arachnida. In: The Natural History of Sokotra and Abd-el-Kuri, Forbes H.O (Ed.). Liverpool Publishing Companies, UK., pp: 175-208.

42:  Proszynski, J., 1966. Remarks on the systematic position pf hemsenattus Iranus roewer 1955 (Arach: Araneae). Sencken- Bergiana Biol. Bd., 47: 463-468.

43:  Proszynski, J., 1992. Salticidae (Araneae) of the old world and pacific islands in several US collections. Ann. Zool. Pan., 44: 87-163.

44:  Rafiinejad, J., S. Tirgari and M. Balali, 2000. Dispersion of poisonous species genus Latrodectus sp. In North of Khorasan and their medical importance (Araneae: Theridiidae). Scientific Res. Q. Shahed Univ., 27: 31-38.

45:  Roewer, C.F., 1955. Die araneae, solifuga und opiliones der sammlungen des herrn Dr. K. Lindberg aus Griechenland, Creta, Anatolien, Iran und Indien. Goteborgs Vetensk Vitterh. Samh. Handl. Bd., 8: 3-47.

46:  Roewer, C.F., 1959. Die araneen der Oesterreichischen Iran expendition 1949/50. Sber. Ost. Akad. Wiss. Abt. I. Bd., 64: 751-782.

47:  Saaristo, M.I. and A.V. Tanasevitch, 1995. Three new Lepthyphantes menge, 1866, from Iran and Turky. Arthropoda Selecta, 4: 61-64.

48:  Senglet, A., 1974. Pholcus nouveaux d'Iran (Araneae: Pholcidae). Revue Suisse Zool., 81: 803-812.

49:  Simon, E., 1893. Etudes arachnologiques. 25e Memoire. XL. Descriptiones d'especes et de generes nouveaux de l'ordre des Araneae. Ann. Soc. Ent., 62: 299-330.

50:  Werner, F., 1936. Reptilien und gliedertiere aus persien. Festschr. Strand. Bd., 2: 193-204.

51:  Wesolowska, W., 1986. A revision of the genus Heliophanus C.L. Koch, 1833 (Aranei: Salticidae). Ann. Zool. PAN., 40: 1-254.

52:  Wunderlich, J., 1995. Revision der Titanoeca tritis-gruppe. mit zwei neubeschreibungen aus der Westlichen palaarktis. Beitr. Araneol. Bd., 4: 731-738.

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