Subscribe Now Subscribe Today
Research Article

Increment Characteristics for Man-Made Stand of Norway Spruce (Picea abies L. Karst) in North of Iran

A. Fallah, H. Jalilvand, M. R. Pormajidian, S. Mohammadpoor Pashakolaei and Y. Kooch
Facebook Twitter Digg Reddit Linkedin StumbleUpon E-mail

In this study, the afforestation stands of 18.1 ha with planting interval 2x2 m of Picea abies, at the age of 44 years old was investigated to compute increment and wood production in Kelardasht plantation (North of Iran). This stand was inventoried using region map and 70x70 m inventory grid. The samples were taken by randomized-systematic method. Increment statistical factors in 32 sample plots with 500 m2 area were measured. Inside each plot, Diameter at Breast Height (DBH) of whole trees and height of four witness trees (two thickness trees inside the plot and two nearest trees to center of plot) were measured. Results of this research showed that survival percentage of Picea abies at the age of 44 years old was 39.3% in Kelardasht region. Also, the average of stand diameter, height, basal area, volume, annual volume increment and form factor were 16.86 cm, 18.20 m, 23.59 m2 ha-1, 193.73 m3 ha-1, 4.4 m3/ha/year and 0.46, respectively. Considering obtaining results and comparing them with its main habitat, the stand increment amount and statistical parameters are low.

Related Articles in ASCI
Similar Articles in this Journal
Search in Google Scholar
View Citation
Report Citation

  How to cite this article:

A. Fallah, H. Jalilvand, M. R. Pormajidian, S. Mohammadpoor Pashakolaei and Y. Kooch, 2009. Increment Characteristics for Man-Made Stand of Norway Spruce (Picea abies L. Karst) in North of Iran. Journal of Applied Sciences, 9: 950-955.

DOI: 10.3923/jas.2009.950.955



Pay attention to, the most species of Iran North forests have low increment, need to woods with tall wood elements (fiber and tracheid) in woody industries and also considering to establishment of forgery big industries in world thus, need to afforestation is increasing day to day. It is look that plantation of species with high increment is able to answering to these needs. With considering, plantation of native species are proposed always, but sometimes because of economical reasons, higher increment and ensuring of appropriate wood for marketing, it is possible to plantation of exotic species (Rezayee, 1995; Khodabakhsh, 1998).

We should culture the species with high increment by arboriculture programming for its utilization in industrial poles. In these conditions conservation of existence species are necessary and plantation with exotic species is proposed by Mosadegh (1996). Investigation of afforestation successfully with Picea abies in plant community of Kelardasht region showed that the parcel with number 4 and with fertility index of 26 is the best site. This species in this habitat with production of 120 m3 ha-1 and age of 25 years old is similar to the best site of Picea abies in European (Mirbadin and Sagheb, 1991).

Pormajidain (1992) investigated increment characteristics for Picea abies in Kelardasht region (North of Iran) by randomized-systematic method with considering age of 29 years old and diameter mean of 11.05. Results his research showed that the diameter, height, volume increment average were 0.38 cm, 0.32 m and 1.56 m3 ha-1, respectively. Rezayee (1995) surveyed Picea abies afforestation stand in age of 35 years old in Lajim region (North of Iran) with plantation distance 2x2 m using random-systematically method. He was resulted that volume, basal area, height and diameter increment average were 4.3 m3/ha/year, 0.4 m2 ha-1, 0.4 m and 0.5 cm, respectively.

Johnssson (1995) investigated the Picea abies stands situated in 55° to 66° latitude and mean age of 41 years old. Results of his research showed that mean of stand density and diameter is 1640 ha-1 and 25 cm, respectively. Also, it is resulted that viability percentage of Picea abies young stands are less in heavy clay soils in compare to the other soil types. The institute of forests and rangelands researches in Iran studied the efforestation successfully of Picea abies stand in Kelardasht region. The obtained report is indicating adaptation of this species with its site (Khodabakhsh, 1998). Gorjy and Gholy (1997) investigated this species in Sangdeh region of Mazandaran province (North of Iran). Their research showed that Picea abies with provenance of Yugoslavian is adapted with site conditions and viability of this species is 95%.

Siahipoor et al. (2000) studied the afforestation 23 years old of Picea abies with plantation distance 2x2 m in Asalem region (North of Iran). The mean of diameter, height, basal area and mean of annual volume increment were recorded 14 cm, 10.5 m, 34.3 m3 ha-1 and 7.5 silve ha-1, respectively. Misson et al. (2002) showed that stand density can change the relation between climate and increment as in single trees; the thinning can increase resistance of tree opposite to drought. Makinen et al. (2002a) mentioned that thinning is due to increasing of radial increment in low sections of tree stem in compare to higher heights.

Makinen et al. (2002b) investigated the changes of radial increment for Picea abies in different latitudes and altitudes in north and center of European. Results of theirs research showed that increment changes in 10 years periods were similar in every site, but had significant different in different sites. Jaakola et al. (2006) studied the Picea abies stand in Finland. Results of his research showed that thinning operations increased the speed of single tree increment but special weight was reduced. Increasing speed of increment is possible to reduce of tracheid length and size of fibers is affected, also. Parn (2006) resulted that climate changes has significant effects on radial increment but pollutions have low effects on radial increment of Picea abies stand.

Raiskila et al. (2006) studied the characteristics of wood in Picea abies species. They reported the mean of annual ring cross about 2.76±1.07 to 3.70±1.22 mm. Mean of special weight in Picea abies wood was 461±0.077 g cm3, also. Zangrele and Oberhuber (2007) studied the changes of radial increment in Picea species. They resulted that felling of aside dominant trees, is due to improvement of tree radial increment. Investigation and growth study and operation of Picea abies afforestation stand and its comparison with origin sites of this species and performed afforestations can be an appropriate field for the better management of aforementioned afforestations. This research was investigated the increment characteristics of Picea abies including diameter, height, basal area and volume in Kelardasht region (North of Iran) and obtained results will compare to results some of other regions.


Study area: Study area (18.1 ha-1 areas) was afforestation with Picea abies (L.) in 1965 by Total Official of Natural Resources-Noushahr (TONRN) (North of Iran) with initial plantation distance of 2x2 m. These areas are situated in forest management plan of Marzan Abad. Study area is located on 45 km southwest far from Chalous town (Kelardasht region) between 36° 29’ 54" to 36° 29’ 52" North latitude and 51° 10’ 51" to 51° 10’ 00" East longitude. The general aspects of these forests are Northern. The average slope of forest field is about 15% (Min. 5% and Max. 27%), the height of forest area at sea level starts from 1250 m and continues till about 1350 m (Mohammadpoor, 2007). This research was performed in the summer of 2006.

Sampling method: Pay attention to evenness of stand and facility in performance, randomized-systematic method was used for accomplishment of this research. Area of sample plots with considering at least 10-15 trees in every sample plot (Zobeiry, 2000) were selected 500 m2 with circle form. Numbers of 32 sample plots with grid dimensions of 70x70 m were designed and inventory intensity was 10.2% in this study that was sufficient pay attention to obtained results by Pormajidian (1992), Keshavars (1992), Khodabakhsh (1998) and Haghi (2002). In every sample plots, diameter at breast high whole trees using caliper and height of witness four trees (two largest diameter and two nearest to center of sample plots) and totally, height of 128 witness trees in whole areas were recorded by means of suunto clinometer’s.

Investigation normality of Picea abies stand: Normality distribution is defined by probability density function (Eq. 1) (Zobeiry, 2000).


μ = Real mean of community
σ = Standard deviation
x = Quantity that calculating of its frequencies is considerable
e = Natural logarithm, that is equal to 2.7183
N = Total No. of community members

Pearson coefficient was used for calculation of skewness (Eq. 2, 3) (Zobeiry, 2000; Rahimnejad, 2002).



b = Pearson skewness coefficient
= Community mean
Md = Medium
Sx = Standard deviation
L = Low limit of class that medium is situated in it
n = No. of data
fc = Density frequency before class that medium is situated in it
fi = Unconditional frequency of class that medium is situated in it
C = Class cross

Calculation of form factor and stand kurtoisis: Form factor and kurtoisis were calculated using Eq. 4 and 5 (Namiranian, 1991; Zobeiry, 2000).



Fn = Kurtoisis coefficient of stand
= Average of height (m)
= Average of diameter at breast height (cm)
f = Form factor of tree
V = Tree volume (m3)
h = Height of tree (m)
g = Basal area at breast height (m2)


Distribution number per hectare and stand normality: The diagram of number distribution in diameter classes and normal curve showed that studied stand has even aged structure (Fig. 1) and probability density function is as below:

For confidence normality and abnormality of stand distribution, Chi-square test was used which calculated 34.601 for this stand. Pay attention to Chi-square test of table at level of 5% and degree freedom 8 (equal to 15.507), the supposition of different between numbers per hectare of Picea abies stand in Kelardasht with normal distribution are significant. Pearson coefficient for distribution number per hectare of stand on basis of diameter classes was calculated –0.76. Aforementioned stand has 982 No. ha-1 and considering initialing plantation distance 2x2 m and number of initial plantation 2500 No. ha-1, natural mortality, cultural inventions, disallowable felling, disease and pests, the viability percentage was 39.3%.

Fig. 1: The normal curve of number distribution per hectare for Picea abies stand

Table 1:

Statistical characteristics for Picea abies in Kelardasht region at 44 years old

Diameter, height, basal area and volume increment mean: In even aged stand that diagram of stand trees distribution in diameter classes is near to normal status, the arithmetic mean of diameter and height are used. Diameter mean, diameter average increment, arithmetic mean of height, average increment of height, mean of basal area, average increment of basal area, growing stock, average increment of volume were calculated 16.86, 0.38 cm, 18.20, 0.41 m, 23.59 m2 ha-1, 0.54 m2/ha/year, 193.73 m3 ha-1 and 4.4 m3/ha/year, respectively (Table 1).

Form factor and kurtoisis of stand: Forma factor calculated 0.46 for Picea abies stand. Also, statistical characteristics including standard deviation, standard error and coefficient variance (%) for form factor of Picea abies were calculated 0.03, 0.01 and 6.5%, respectively. Kurtoisis coefficient is one of the most factors that are used for judgment in relation to stand stability. This coefficient was 107.95% for studied stand (Fig. 2).


Distribution number per hectare and stand normality: The curve of number per hectare in different diameter classes (Fig. 1) showed that investigated stand has even aged structure. Chi-square test showed that the curve of number per hectare is not similar to normal distribution curve. Skewness of stand is high, also (Rahimnejad, 2002). Number per hectare was more than normal status at diameter classes of 12-14 cm that should be considered in cultural operations. Viability percentage and number per hectare of Picea abies stand were compared pay attention to this study and the other performed studies (Table 2). Table 2 showed that Picea abies stand of Kelardasht region has viability percentage and number per hectare more appropriate with considering to stand age in compare to the other mentioned sites.

Diameter, height, basal area and volume increment average: Pay attention to performed studies (Mirbadin and Sagheb, 1991; Pormajidian, 1992; Rezayee, 1995; Siahipoor et al., 2000; Dohrenbusch et al., 2002; Siahipoor et al., 2002), it is mentionable that diameter increment of this stand had not high changes from 29 years old up to now. Also, diameter increment average is lower than sites with three grades of French (Table 3). Because of number per hectare of low diameter classes are high in stand, therefore, trees have competitive status one another and the stand has higher height increment in compare to diameter increment.

For improvement of stand diameter increment, harvesting of low diameter classes should be considered in programming of cultural operation in order to increasing of stand diameter increment.

Table 2: Comparison of viability percentage of Picea abies in different regions

Also, with considering Table 3, it is clear that height increment average of stand is equal to French sites, relatively. Because of existence high density and increasing of competition in stand, the diameter increment is reduced and height increment will increase, thus, stand stability will decrease.

Basal area increment of stand are lower than to the other sites similar to diameter increment because of high density of low diameter class in stand (Table 3), which is due to reduction of diameter, basal area and volume increments. Pay attention to performed studies (Assman, 1961; Pormajidian, 1992; Rezayee, 1995; Dohrenbusch et al., 2002; Siahipoor et al., 2002), it is considerable that volume increment of stand was lower than to different sites in Iran and out of Iran and also has lower growing stock (Table 4) and performing of appropriate culture operations is necessary for its enhance.

In order to increase of production in area unit and improvement of quantity and quality characteristics of Picea abies afforestation in Kelardasht region, the operation of thinning should be performed with high precession pay attention to site typical conditions, high kurtoisis coefficient and skewness coefficient of stand. Because of, every intervention without consideration to stand conditions is due to reduction of tree resistance opposite to environmental parameters such as wind and snow, etc. and it is increased fall of stand trees by wind.

Form factor and kurtoisis of stand: Form factor of stand is depending to some of factors such as age, site and species, even aged or uneven aged of stand and cultural method of forest. Also, form factor is more in young stands and lower in stands with high ages (Zobeiry, 2000). Form factor mean of Picea abies stand in Kelardasht region calculated 0.46 that is appropriate, almost with considering the low age of stand. Stability coefficient was calculated 107.95% for Picea abies stand in Kelardasht stand at age of 44 years old, therefore, the stand sorted in very instability class (Namiranian, 1991).

The curve of stability coefficient on basis of diameter classes is showing that stability coefficient has descending status and the stability of stand will increase with increasing of diameter (Fig. 2).

Table 3: Comparison of diameter, height and basal area increment for Picea abies in different regions

Table 4: Comparison of volume increment for Picea abies in different regions

Fig. 2: Scatter plot and the curve of stand kurtoisis

High kurtoisis coefficient of this stand is due to stand density, competition between species for achievement to light and laying in high stair and do not thinning of stand in appropriate time. With appropriate cultural operation and thinning of stand, it is possible to reduce this coefficient but, thinning operation should be performed with high precession because of stand is very instability and pay attention to superficial roots of Picea abies species, the danger of fall by wind is very high in stand. Siahipoor et al. (2002) estimated the stability coefficient for Picea abies at 27 years old about 89 and 78% for Oroston and Pisesoon regions (North of Iran), respectively. Therefore, Oroston Picea stand and Pisesoon Picea stand are sorted in instability and stability classes, respectively.


Mean number per hectare of Picea abies in study area was 982 and almost 60% of initial planted sapling were deleted by reason of competition, primary harvesting and the other environmental factors such as snow, wind and disallowable felling and the stand missed its normal status and has high density in low diameter classes. Mean of diameter, height, basal area and volume increments in study area and its compare with the other sites is indicating low increment and yield in stand. Probably, low increment in surveyed stand is due to disallowable felling and do not performing of appropriate cultural operations in stand that is due to stand distance of normal status.

1:  Assman, E., 1961. The Principles of Forest Yield Study. 1st Edn., Pergamon Press, Oxford, New York, ISBN: 91-576-6069-7, pp: 506-506.

2:  Dohrenbusch, A., S. Jaehne, M. Bredemeire and N. Lamersdorf, 2002. Growth and fructification of a Norway spruce (Picea abies) forest ecosystem under changed nutrient and water input. Ann. For. Sci., 59: 359-368.
CrossRef  |  Direct Link  |  

3:  Gorjy, B.Y. and Z.M. Gholy, 1997. Investigation and comparison successfully of planting with Vase and bare root forms in autumn and spring seasons in Sangdeh region (North of Iran). Paj. Saz. J., 37: 60-61.

4:  Haghi, A., 2002. Estimation increment of Cupresus horizontal in natural and artificial stands by using of stem analysis in Guilan province. M.Sc. Thesis. Guilan University, pp: 101.

5:  Jaakola, T., H. Makinen and P. Sarapaa, 2006. Tracheid properties of Norway spruce (Picea abies) grown at long-term fertilization thinning regions. WSE, Stockholm, T.J., METLA. Finnish Forest Research Institute, pp: 30-31.

6:  Johnssson, T., 1995. Site index curves for Norway spruce plantations on farmland with different soil types. Studia. Forestalia. Suecica., 198: 19-19.

7:  Keshavars, G.H., 1992. Investigation of pinus teada L. in Talesh region of Guilan province. M.Sc. Thesis. Tehran University, pp: 112.

8:  Khodabakhsh, S., 1998. Survey quantitative and qualitative of picea abies in pure and mixed stands of picea artificial forest in Kelardasht region. M.Sc. Thesis. Tehran University, pp: 137 (In Persian).

9:  Makinen, H., P. Nojd and A. Isomaki, 2002. Radial, height and volume increment variation in Picea abies stands with varying thinning intensities. Scand. J. For. Res., 17: 304-316.
Direct Link  |  

10:  Makinen, H., P. Nojd, H.P. Kahle, U. Neumann and K. Tveite et al., 2002. Radial growth variation of Norway spruce (Picea abies) across latitudinal and altitudinal gradients in central and northern Europe. For. Ecol. Manage., 171: 243-259.
CrossRef  |  Direct Link  |  

11:  Mirbadin, A. and T.K. Sagheb, 1991. Afforestation successfully with Picea abies in different communities of Kelardasht region. Jang. Sen. J., 75: 36-75.

12:  Misson, L., C. Vincke and L. Devillez, 2002. Frequency responses of radial growth series after different thinning intensities in Norway spruce (Picea abies) stands. Forest Ecol. Manage., 6050: 1-13.

13:  Mohammadpoor, P.S., 2007. Investigation of diameter growth of Picea abies using stem analysis (Case study: Kelardasht region-Mazandaran province). M.Sc. Thesis. Mazandaran University, pp: 136.

14:  Mosadegh, A., 1996. Silviculture. 1st Edn. Tehran University Press, Iran, pp: 481.

15:  Namiranian, M., 1991. Investigation of increment models application in forest. Nat. Res. J., 45: 93-102.

16:  Parn, H., 2006. Radial growth of conifers in regions of different cement dust loads. Proc. Estonian Acad. Sci. Biol. Ecol., 55: 108-122.
Direct Link  |  

17:  Pormajidian, M.R., 1992. Researches in relation to results of afforestation with Picea abies in Kelardasht region. M.Sc. Thesis, Tehran University, pp: 162.

18:  Rahimnejad, S., 2002. Quantitative investigation and estimation of Pinus teada volume in Lakan region of Guilan. M.Sc. Thesis, Guilan University, pp: 80.

19:  Raiskila, S., P. Saranpaa, K. Fagerstedt, T. Laasko and M. Loija et al., 2006. Fast growth rate reduces wood density and strength properties of Norway spruce. Growth rate and wood properties of Norway spruce cutting clones on different sites. Silva Fennica., 40: 247-256.
Direct Link  |  

20:  Rezayee, A., 1995. Quantitative and qualitative investigation of Picae abies afforestation in Lajim region. M.Sc. Thesis. Gorgan University, pp: 106.

21:  Siahipoor, Z., Z.V. Aman and T.K. Sagheb, 2000. Investigation of increment and wood production of Picea abies in Asalem region. Paj. Saz. J., 46: 64-67.

22:  Siahipour, Z., T. Rostami, T.K. Sagheb, 2002. Investigation of sustainable Picea abies in forestations of Guilan province. Jan. Sen. J., 312: 1-53.

23:  Zangrele, P. and W. Oberhuber, 2007. Dendroecological analysis of the impact of debris flows on a high mountain forest ecosystem: A case study in Northern Limestone Slops (Tyrol, Austria). European Geosciences Union, Geophysical Research Abstracts, Vol. 9: 02134.

24:  Zobeiry, M., 2000. Forest Inventory (Measurement of Tree and Stand). 1st Edn., Tehran University Publisher, Iran, ISBN: 964-03-4524-5.

©  2021 Science Alert. All Rights Reserved