Abstract: Stem anatomy and nodal vasculature are studied in seven Vicia species. The stem anatomy shows the presence of 30 characters proved to be taxonomically useful, of these: outline of cross section, number of its sides, presence of brown tanniniferous cells in the epidermis, presence of cortical fibre strands, number of vascular bundles in the ring and pith cell shape. Only the epidermis of V. sativa is characterized by the presence of some brown tanniniferous cells and the cortex of V. faba is characterized by the presence of four cortical fibre strands, two above the two polar bundles and two above the two cortical bundles in the wings. At the beginning of nodal vasculature, the number of the vascular bundles increases from ten to fourteen in V. sativa, V. ervilia, V. monantha and V. villosa, from nine to thirteen in V. peregrina, from nine to twelve in V. narbonensis, or from 18-20 in V. faba. The studied Vicia species are characterized by an unusual type of stele in which the two cortical bundles divide each into two bundles; one supplies the stipules and the other enter the leaf base. The median petiole trace divides into three leaving the main stele and passes out to the petiole. Each lateral branch of this median trace divides again and the outermost bundle fuses with the incoming lateral petiole trace from cortical bundles supply. The five bundles in the petiole compose of the lower most three are entirely median in derivation, while the two uppermost bundles are composite. Thus the lateral leaf traces are derived from the two cortical bundles of the lower internode below the insertion of their leaf, beside to the two lateral bundles deriving from the median leaf trace and the stipules are supplied by the two cortical vascular bundles only and not by the main vascular ring. A complete replacement occur for the two cortical bundles, they substitute at each node by two new bundles originating from the two lateral bundles of the main ring opposite to them supplying the stipules of the next node. Axillary bud is supplied from the two lateral bundles on either side of the median leaf gap.